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The West that Was . . . No Longer Is

Losing Ground

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Precipitation - A Driving Force

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Elk and Mule Deer Interactions

Mule Deer Regions - No Two are Alike

Plant Communities in Trouble . .

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Mule Deer Regions - No Two are Alike


Map of Mule Deer Ecoregions in the West

No two mule deer populations are alike because where they live differs from one part of their range to another. Biologists refer to the different areas as “ecoregions”. By studying each ecoregion, biologists can better identify the factors that are limiting the growth of mule deer populations, and predict responses by mule deer populations to changes in habitat. Habitat quality has an effect on survival of fawns, the most important factor in determining how well a population fares from season to season, and year to year.

Biologists have identified seven ecoregions that mule deer call home. Each ecoregion is briefly summarized including a physical description, a description of the deer, the climate, limiting factors that reduce the productivity of deer, and recommendations to improve mule deer populations.

All ecoregions are subject to the limiting factors such as urbanization, fire suppression and drought described in this publication. The limiting factors listed for each ecoregion in this article are some of the most important, but certainly not the only factors, limiting mule deer populations.

Although each region is ecologically different, some common factors exist:

- Generally, habitats conditions that are less productive for mule deer.

- Human caused factor such as fragmentation of habitat, changes in fire regimes, livestock management and changes in plant communities have limited deer populations.

- Return to higher mule deer numbers will require stronger land use planning and restoration efforts on a large scale.

- Climate and weather play an important role in habitat quality in each region.


Coastal Rain Forest Ecoregion

Description: Along the west coast of North America from northern California through southeast Alaska. Known for its dense rain forests of western hemlock, Sitka spruce and natural and commercial forests of Douglas fir. Clearcutting is common in commercial forests, and provides excellent habitat for mule deer for eight to 10 years after harvest when grass, forbs, shrubs and saplings are common. In the northern part of this region where winter snowfall can be heavy, it is important to retain stands of mature trees to intercept the snow.

Mixed conifer habitat of Coastal Rainforest Ecoregion, west slope of Cascade Range, Douglas County, Oregon.  ByTom Keegan.Climate: A marine climate with cloudy days, cool temperatures, high precipitation from fall to spring, and a short, dry summer season. Precipitation ranges from 25 to 120 inches. Soils are coarse and nitrogen-poor.

The deer: Black-tailed deer are the dominant subspecies of mule deer. The deer are primarily nonmigratory, and are well distributed and occur at the greatest densities in early successional habitats in the central and southern part of the region. In the northern part of this ecoregion, deer numbers are greatest on coastal islands, where marine weather lessens the severity of winter. In the far north, winter snow may force deer to lower elevations. Black-tailed deer are often unable to meet their nutritional requirements year round. Fawns rarely breed, and pregnancy rates for yearlings vary greatly from year to year, but are generally low. Deer in this region tend to be older than in other regions because the amount of secure cover deer find in the dense forest limits hunter success.

Limiting Factor: The quality of the plants. Heavy rainfall and soils poor in nitrogen cause nutrients to leach from the soil, and plants have more moisture. Pound for pound, a deer consumes less nutrients while foraging in coastal rain forest than in other regions.

Recommendations:

1. Create more grass, forb, shrub and sapling communities to improve food quality.

2. Maintain forest canopies in places where snowfall is heavy.

3. Manage forests for high quality plant foods to allow for large harvest of deer to reduce overwintering populations, and thus reduce browsing on young conifers.

4. Survey for diseases and parasites.

5. Plant mast producing species such as oak in dry and southern areas.

6. Conduct small, cool controlled burns.


California Woodland Chaparral

Description: Includes the Coast Range of southern California, and lower elevations of the west slope of the Sierra Nevada east into central Arizona.

Live oak-chaparral woodland habitat shot taken of Bloomfield Ranch, Kern County, California. By Marc Hoshovsky.Climate: Hot dry summers, mild wet winters, and periodic droughts create annual grasses and forbs in communities of oak woodland and chaparral. Precipitation ranges from 8 to 30 inches a year. Chaparral was once maintained by frequent, cool fires, but fire suppression created older stands of chaparral with poor quality forage.

The deer: Mule deer populations in this region do not migrate, except for those at higher elevations in the Sierra Nevada and San Gorgonio Mountains. Deer densities are greatest in the northern part of this ecoregion. Nonmigratory deer move in response to changes in habitat on north and south facing slopes.

Limiting factors: Fire. Most of the mule deer range in this region is in private ownership, and fire suppression is a high priority for residents. This region is in a fireadapted habitat, and frequent fires occurred before European settlement. Frequent fires rejuvenate the habitat and improve forage for mule deer. Fire suppression results in infrequent, large, hot fires. The lack of fire results in older, less nutritious plants for mule deer. Weather. Summer and early fall is a difficult time for mule deer because of little rainfall, and dry plants with little nutritional value. Nursing does need high quality forage to nurse fawns and build body reserves for the coming winter.

Recommendations:

1. Use fire to stimulate sprouts of shrubs over a large landscape.

2. Stimulate new growth of desired plants using light livestock grazing.

3. Minimize effects of livestock along streams and uplands to improve forage for mule deer on fall and winter ranges.


Southwest Deserts Ecoregion

Description: Includes the southern portions of California, Arizona, New Mexico and west Texas, extending into northern Mexico.

Spring wildflowers in the Southwest Deserts ecoregion. By Arizona Game and Fish Department.Climate: This region is arid to semi-arid, and has extreme temperatures, high evaporation rates, low rainfall that varies greatly from year to year, periodic droughts and poor soils. Precipitation ranges from 5 to 20 inches annually.

The deer: Deer are nonmigratory and greatly affected by droughts. Fawn recruitment is variable depending on amount and timing of rainfall. During dry years, fawn recruitment is typically below what is needed to maintain the population.

Limiting factors: Rainfall and competition with livestock. Winter rainfall affects the diversity, quality and quantity of next years' spring forbs, which directly affects the number of young deer that are born and survive to adulthood. Winter precipitation stimulates plant growth in the spring. Forbs are critical to the survival of deer in this ecoregion because browse plants don't contain adequate amounts of nutrients. Competition with other forb and grass-eating species such as livestock can have a great effect on mule deer, especially during years when rainfall is limited and range resources are scarce. Overgrazing in drought years can have long-lasting effects.

Recommendations:

1. Create sources of water in areas where water is limiting and where other potentially limiting factors are being addressed.

2. Monitor grazing so that livestock do not remove large amount of plants, particularly in years where drought or other climatic conditions stress deer.

3. Work with landowners to provide hunter access to public land.

4. Monitor human sprawl.


Great Plains Ecoregion

Description: The largest grassland ecosystem in North America, extending from central Canada to the Texas panhandle, west to the Rocky Mountains. The region includes a transition from tallgrass to shortgrass prairie.

Mule deer habitat used in the Great Plains ecoregion, Scotts Bluff National Monument in western Nebraska. By Mike Cox.Climate: This region is semiarid, annual precipitation varies between 10 and 33 inches, and temperature varies greatly.

The deer: Mule deer in this region are nonmigratory, although they shift their home range in response to local moisture conditions that affect plant quality. Mule deer forage on agricultural plantings in areas that are irrigated.

Limiting factor: Cover. Drought and severe winter snows can affect mule deer populations. Fire is important in maintaining grasslands. Draws that contain shrubs, hardwoods and moisture provide mule deer with critical habitat, especially in the winter. Grassland and shrub/grassland communities interspersed with draws provide critical year-round habitat for mule deer. Irrigated fields grow nutritious grasses for mule deer forage. Human activities are a doubleedged sword for mule deer. While agriculture provides watering holes and alternative food sources for mule deer, overgrazing by livestock is harmful to the woody draws that provide cover and moisture.

Recommendations:

1. Work with landowners to minimize the effects of severe weather conditions by providing hard woody cover for mule deer by improving grazing strategies and riparian habitats.

2. Provide hunting opportunities consistent with habitat conditions and deer populations.


Colorado Plateau Shrubland and Forest Ecoregion

Description: High elevation areas in western Colorado, eastern Utah, southern Wyoming, and northern Arizona and New Mexico. Habitat ranges from spruce trees at high elevations, ponderosa pine and Douglas fir at mid-elevations, and sagebrush and pinyon-juniper at lower elevations.

Aspen stand in poor condition because there is no aspen generation and the stand is being invaded  by conifers. Fire is needed to restore and rejuvenate this stand. By Dan Stroud.Climate: Much of this region is above 5,000 feet and includes many mountain peaks above 15,000 feet. Precipitation ranges from 8 to 24 inches. Winters can be severe.

The deer: Deer are migratory because of the heavy winter snowfalls at high elevations. Deer populations are most affected by severe winters that cause nutritional stress, high fawn mortality and lower fawn recruitment. Some lower elevation ranges can be summer range limited. Livestock grazing may affect the quality of forage available to deer.

Limiting factors: Severe winters and droughts can impact the productivity of mule deer by causing high fawn mortality. Improper livestock grazing has caused changes in mule deer winter range.

Recommendations:

1. Limit disturbance to existing winter range, and acquire additional winter range.

2. Improve quality and quantity of winter range habitat.

3. Maintain stands of aspen for mule deer fawns and summer range.

4. Limit development of and disturbance to summer range in areas where summer range is limiting.


Intermountain West Ecoregion

Description: The mountain ranges west of the Rockies, east of the Sierra Nevada, north of Colorado and south of Canada. The Great Basin, a large semiarid basin, makes up a big part of this land mass. Pinyon-juniper woodlands, conifer forests and aspen woodlands are common at higher elevations.

Steens Mountain, Oregon in the Intermountain Ecoregion. By Tom Keegan.Climate: Lower elevation communities receive less than 12 inches of precipitation a year. Areas to the north and at higher elevations receive most of their precipitation as snow.

The deer: If you could draw a bull’s-eye around the portion of the West that was once the center of mule deer distribution, you would draw it around this region. Mule deer typically migrate in this region (although some do not), spending summer in conifer forests at higher elevations and winter in lower elevations. Deer densities are highest in places where vegetation and topography are diverse. Agriculture and urban development have hurt mule deer populations in this region by destroying shrub communities and reducing winter range.

Limiting factors: Competition with livestock, agriculture, urban development and timber management. Each year, thousands of acres of sagebrush habitat and valleys are being overtaken by pinyon- juniper stands, much to the detriment of mule deer. In the southern part of the region, invasive plants such as cheatgrass and changes in fire cycles are limiting mule deer productivity. Habitat in spring and summer affect mule deer productivity more than severe winters because the quality of spring and summer range affects the number of fawns surviving to adulthood. Urban development may affect recruitment because it is occurring in mule deer winter range.

Recommendations:

1. Manage motorized traffic to benefit mule deer.

2. Manage forests for both early and late successional stages to meet year-round needs of mule deer.

3. Protect and plant important browse species for mule deer, especially in winter ranges.

4. Manage wildfires on mule deer ranges to avoid cheatgrass invasion.

5. Manage livestock grazing to minimize impacts to mule deer along streams and in aspen habitats.

6. Develop cost-effective ways to reduce pinyon-juniper invasion, and place a priority on developing a patchwork of habitats so that mule deer have woody cover near places to forage.


Northern Forest

Description: The higher elevations of the Cascades and Sierra Nevadas in the three most western states, as well as northern Idaho, western Montana and Wyoming, northern Washington, and the western Canadian provinces. Pine, spruce, fir, Douglas fir and larch are the dominant forest types, and forests become more thin as elevation increases. Mule deer are not found very far north of the northern boreal forest in subarctic woodlands.

Male black-tailed deer in mixed conifer forest, Douglas County, Oregon, December 1998. By Tom Keegan.Climate: Winters are long and cold. Average annual precipitation varies with elevation and topography, from 10 inches to as much as 120 inches.

The deer: Because of severe winters and heavy snowfall, most of the deer in this region are migratory, although some are yearround residents at lower elevations. The growing season is short, and the quality of food mule deer find during this critical time is high. Deer follow retreating snow in search of food.

Limiting factors: Severe winters. Deer follow the growth of plants throughout the growing season. It is only when severe winters and deep snow limit their ability to forage that they experience die-offs and high mortality. If mule deer populations experience a die-off, there is excellent chance for recovery as a result of spring and summer habitat conditions. The greatest threats to deer in this region are development and disturbance of winter range, and barriers to migration.

Recommendations:

1. Acquire winter range habitat and minimize housing developments to protect and enhance winter ranges.

2. Use fire to maintain shrubdominated habitats.

3. Maintain forest shrubs, forbs, grasses and saplings to provide foraging habitat in spring, summer and fall.

4. Avoid and manage forest encroachment into high elevation meadows.

5. Avoid barriers to migration.

6. Manage deer populations based on the ability of winter range to support them, and avoid overharvest in years when early winters send migratory deer to lower elevations.


Mule Deer, Changing Landscapes, Changing Perspectives, is a series of non-technical articles based on technical papers from the book, “Mule Deer Conservation: Issues and Management Strategies” Published by The Berryman Institute, Utah State University.

The contents of this web page may be photocopied or reprinted for noncommercial purposes using the citation listed below:

Mule Deer Working Group. 2003. Mule Deer: Changing landscapes, changing perspectives. Mule Deer Working Group, Western Association of Fish and Wildlife Agencies.