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WAFWA Mule Deer Working Group

What's in a Name?

Of Shipwrecks and Captives

The West that Was . . . No Longer Is

Losing Ground

The Mounting Pressure of Development

A Place for Predators

Precipitation - A Driving Force

Wilderness Breakup

Elk and Mule Deer Interactions

Mule Deer Regions - No Two are Alike

Plant Communities in Trouble . .

Mule Deer Diseases

Supplemental Feeding - Just Say No

Learning By Doing

Managing Deer Herds with Harvest

Our Summary

WAFWA

Losing Ground

Fire, invasive species and livestock management have changed western landscapes

Fire

Of all the factors that have shaped the ecoregions in which mule deer exist, fire has been the strongest one with the greatest positive influence. Fire is a critical force in maintaining and creating habitat for mule deer because fire sets back succession.

Succession is the orderly and progressive replacement of one plant community by another until a fairly stable community occupies an area. If left alone, an abandoned crop field will not remain in that state for very long. Generally, grasses, forbs and weeds will begin to grow, followed by brushy plants, then by saplings that invade open areas, until the site is finally occupied by a stand of trees. Historically, fire has been the most effective tool in maintaining grasslands across the United States. Today, it is still considered to be the most important tool a biologist has to manage habitat.

A quick peak at national historical wildfire data provides insight into the frustration land managers face with fire suppression efforts.

Decade # of acres burned
1920s 26 million acres
1930s 39 million acres
1940s 23 million acres
1950s 9 million acres
1960s 4 million acres
1970s 3 million acres
1980s 4 million acres
1990s 3.6 million acres

Since the decade of the 1940s, fires have not burned in double digit numbers in the United States. Wyoming Game and Fish Biologist Steve Kilpatrick attributes the heyday populations of mule deer in the middle of the 20th century to the quantity and types of fires that burned in the decades prior to the 1950s.

“In the 1920s through 1950, we had some massive burns and resprouting shrubs,” said Kilpatrick. “We had high quality browse, and a lot of quantity - lots of acres of good (mule deer) groceries. Browse was nutritious, young, palatable and easy to digest.” Now the plants are older, and when it rains or they are browsed, they don’t respond as vigorously.

Forest fire. By Gary Schafer.In addition to fewer fires burning on fewer acres, fire suppression has changed the intensity and rate at which fires burn, resulting in different and unpredictable communities of plants. Fuel loads build up such that when infrequent fires occur, they cover large amounts of land and burn very hot. A recent example of that is the Rodeo fire near Pinedale, Arizona, that burned at 2,000 degrees Fahrenheit. In general, areas that burn fast and hot become monocultures, in which there are fewer types of plants that are similar in age.

Daryl Lutz, Wildlife Management Coordinator for Wyoming Game and Fish in Casper, said that lack of fire is creating overaged less useful aspen and sagebrush stands.

“On mule deer summer range, aspen communities are being lost at an alarming rate due to natural vegetative succession,” said Lutz. The reason these habitats are being lost is lack of fire.

“In an aspen stand, you can see a vegetative response within one or two years of a fire,” said Lutz. But some plant communities do not respond as quickly.

Lutz said, “In a sagebrush stand, it could be up to twenty years.” Lutz emphasized the importance of creating patterns of habitat.

“Whenever we do things in sagebrush communities, we always emphasize and tailor our prescriptions to a mosaic of burned and unburned,” said Lutz. “We’re starting to evaluate how we should be doing prescribed burning so we don’t eliminate brood rearing or nesting habitat for sage grouse, and help other species.”

Ken Mayer of the California Department of Fish and Game emphasized the changes that happen to a landscape over time if small, cool, frequent fires are replaced by large, hot, infrequent fires.

"Hot fires burn minerals from the soil, and you don't get the regeneration you should," said Mayer. This lessens the potential of the site to be productive, and ultimately results in long-term changes to the habitat. "In alpine communities, there is about a three month growing season," said Mayer.

"In some of that country, the snow doesn't come off until July. The plants have a short window to grow, and have adapted to fires over a long period of time. If you eliminate fire, then introduce fire in a big way (a large, hot, intense fire), it takes 10 years for those plants to become useful for mule deer again."

Kilpatrick echoed the consequences of large, hot fires. “Mother Nature says you can pay me now or you can pay me later with interest,” said Kilpatrick. “Suppressed fires will be a lot larger, and the intensity and severity will be greater when they do burn. Wildlife love resprouting shrubs. But fires that burn hot can kill resprouting species of shrubs. It’s quite a while before the moonscape appearance disappears. We’re exacerbating the situation by our actions.”

Mule deer thrive in early successional habitats, where forbs, grassy plants and shrubs dominate. These environments are not as stable as forest habitats, and they rely on fire or some other type of disturbance to return them to an early successional stage. If they are not disturbed, they eventually become more stable plant communities dominated by trees and large shrubs. Tree-dominated habitats offer mule deer a place to retreat from severe weather, but these areas offer very little in the way of food. That is why it is important to provide mule deer with a mosaic or pattern of habitats that can provide food, cover and water.

Arizona Game and Fish Department’s Chief of Research Jim deVos said large, hot fires contribute to soil erosion.

“Another problem associated with the catastrophic fires that are occurring due to long-term fire suppression is that virtually all of the vegetation is lost, which increases soil erosion,” said deVos. “It is important to remember that it took eons to build the top soil layer, and its loss will alter the lands’ ability to rebuild. Where this occurs, the land may never recover its capacity to support wildlife populations as it did before these incredibly intense fires.”

To complicate matters, habitats with plant species such as mountain big sagebrush are experiencing fires every 100 or more years compared to pre-European settlement fire frequencies of 12-25 years. Wyoming big sagebrush, a habitat with large amounts of the invasive plant, cheatgrass, is now subjected to fires every 10 years instead of every 50 to 100 years. Drastic changes in fire frequency may result in changes in the types of plants found in a given area.

Kilpatrick said that drought years compound the problem, making it more difficult for biologists to use prescribed fire.

“We’re using prescribed fire as much as we can, but it’s more difficult to use during these drought years because of the risk factor,” said Kilpatrick. “It takes someone that can find the dotted line to say they’ll be responsible for doing prescribed burns in a risky situation. But fires normally burn twice as many acres in drought years.”

Kilpatrick said land managers are behind the curve burning on a landscape scale, especially compared to the amount of land that used to burn on an annual basis.

Kilpatrick said federal agencies responded to the Yellowstone Fire in the late 1980s with a strong educational effort, but that habitat change often occurs over the longterm, oftentimes longer than the life span of a human being.

“Fire was THE main player forming the very landscape that we cherish and want to protect now,” said Kilpatrick. “People realize it’s a dynamic system, be it ever so slow. For example, aspen needs a fire every 80-100 years. People don’t see those changes taking place in their lifetime. But the public is accepting fire – they just don’t want to see their homes burned down.”

To avoid seeing homes burned, people are willing to pay a steep price. Suppression costs for wildfires are easily three to five times greater than the cost of prescribed fire per acre. In the last seven years, the cost of fire suppression for federal agencies has ranged from a low of $256 million in 1997 to a high of $1.36 billion in 2000.

According to Kilpatrick, the effects of fire suppression are worsened because of habitat fragmentation.

“You couple what has happened with fire, and compound winter range being used by urban sprawl, and then our exploration and development for oil and gas on winter ranges – it’s fragmenting habitat,” said Kilpatrick.

Burned sagebrush habitat from a wildfire that has regrown into exotic plant cover of mustard and cheatgrass in the foreground. The area in the middle of the picture was disced and planted with a mixture of native and nonnative seeds. The dark green hills in the background did not burn and contain native sagebrush. By Ken Gray.As a result of these interactions, Kilpatrick said emphasis should be placed on maintaining critical areas such as important winter ranges.

“We need to put as much management effort on important winter ranges, keep them unfragmented in terms of oil development and maintain high quality forages,” said Kilpatrick.

Kilpatrick attributes increases in predation with habitat fragmentation.

“When you fragment the habitat, prey doesn’t have the landscape to escape predators. Predators have a much easier time catching prey in reducedand fragmented habitat.” He said the increased presence of predators such as mountain lions in suburbs is likely a direct result of the wildlife habitat that has been created at the urban interface.

"We’re so far behind the curve in terms of a landscape scale that we’ll never catch up with prescribed fire,” said Kilpatrick. “Nature will catch us up, as she has during a drought.”

Kilpatrick cautioned, “Don’t blame the predators. They’re the symptom, we’re the problem.”

Invasive Species – A growing threat

What harms 15 percent of our country's ecosystems, costs the United States at least $137 billion a year in lost profit and eradication efforts, and includes a group of about 7,000 species? The answer – invasive species. While many are found throughout the West, one of the most harmful is the winter annual grass called cheatgrass, alias downy brome.

Cheatgrass found its way to the United States from Europe and Asia in the late 1800s on the backs of livestock, and in some grain and hay feed. By 1920, it firmly established itself as a formidable invasive plant.

It is a plant species with few endearing qualities. Cheatgrass is not very nutritious or palatable to livestock and wildlife, although livestock will graze on it in some desert habitats in the winter and spring, and mule deer will browse on it in early spring.

When cheatgrass is present, livestock overgraze native plants, causing direct competition with mule deer for food. But that's not the worst of it. Overgrazing by livestock actually helps cheatgrass gain a foothold, both on the overgrazed land, and on nearby land where invasive plants may not have existed.

What gives cheatgrass the ability to outcompete native plants? John Grahame and Thomas Sisk, editors of "Canyons, cultures and environmental change: An introduction to the land-use history of the Colorado Plateau," a publication from the Center for Environmental Sciences and Education at Northern Arizona University, describe the unique ability of cheatgrass to outcompete native plants.

"Most native bunchgrasses of the Colorado Plateau are perennial, whereas annual plants like cheatgrass grow from a seed, then flower, set seed, and die every year. Cheatgrass usually germinates in fall and grows during winter, opposite the cycle followed by common native perennial grasses. By the time the rain stops in spring, cheatgrass already is maturing its seeds. Unlike native bunchgrasses, cheatgrass then dies by the end of July, avoiding the hottest and driest part of summer.

"Dead cheatgrass burns easily, causing early and abundant wildfires which tend to damage or kill native grasses. During a fire, earlymaturing cheatgrass seeds can take advantage of many nutrients the fire releases to grow large and produce abundant seed, over a thousand per plant in some cases.

"Because cheatgrass quickly develops a large root system in the spring, by the time native grass seedlings start to grow in April or May, cheatgrass has stolen most water out of the top foot of soil. Although mature native grasses can get water from lower soil regions, seedlings cannot get their roots deep enough into soil to access water before drought sets in, and thus, die of thirst. Without this ability to reproduce, native grasses inevitably decline, and so over time, cheatgrass becomes more and more common until eventually it dominates. Cheatgrass often opens the way for secondary invaders such as knapweed and thistle."

A strong invader like cheatgrass poses two threats to mule deer. Cheatgrass outcompetes native perennial forage, and increases the frequency and intensity of wildfires, altering the quality of sagebrush habitat.

When cheatgrass takes hold, it can ultimately outcompete every native plant, creating a monoculture, or a stand of plants that contain one or a few species. Except for the brief period in spring when new green shoots of cheatgrass emerge from the soil, stands of solid cheatgrass have about as much benefit to mule deer as a paved parking lot.

Cal McCluskey, Bureau of Land Management (BLM) Senior Wildlife Specialist, said cheatgrass is altering large tracts of land in the West.

Vast stands of cheatgrass cause frequent, large fires, much to the detriment of mule deer habitat. Intense frequent fires destroy native shrubs such as antelope bitterbrush, an important food for mule deer.

What is being done about the continued threat of invading plants?

The BLM is taking aggressive steps to learn more about the spread of invasive plants and large-scale landscape changes that have occurred since European settlement. And they're developing new and different approaches to combat nonnatives.

"Cheatgrass has created a fire cycle that has altered substantially the historical fire cycle,” said McCluskey. “It has increased the frequency of fires, and in many areas, once the landscape gets burned two or three times, it often comes back in a monoculture of cheatgrass, just prime for burning again and again."

McCluskey said the Columbia River Basin and Great Basin are of particular concern to the BLM.

"Those are the two ecoregions that have had the greatest invasion problems with cheatgrass and other annuals like Medusahead," said McCluskey.

McCluskey said an effort is underway in the BLM to get a better handle on the extent of change on the landscape within the sagebrush ecosystem.

"We have a major effort underway to look at sagebrush habitat throughout its range, but particularly as it relates to sage grouse," said McCluskey. “We're working with the United States Geological Survey to put together a map of sagebrush for the entire West to show current versus historical distribution."

Other large-scale efforts are underway, as well. The BLM is leading the Great Basin Restoration Initiative, an interagency effort to restore areas burned by fire. Reseeding and changing grazing practices are two possible restoration strategies.

"The catalyst for that initiative was the bad fire year of 1991 in the Intermountain West," said McCluskey. "Several million acres burned, some of which was significant mule deer winter range."

Through the Great Basin Restoration Initiative, the BLM in Nevada is cooperating with state and local agencies and nonprofit organizations to restore and maintain native plant communities, and slow or stop the spread of invasive plants on about 10 million acres of public land.

McCluskey also talked about the BLM being the biggest landowner of sagebrush habitat in the West, and the importance of using fire to manage sagebrush.

"We have a major fire rehabilitation program to go back into areas that burn and reseed," said McCluskey. "State wildlife agencies provide the native seed and bare root stock. We target the areas with the highest probability of success."

Reducing the size and frequency of fires by creating fire barriers such as green strips is another effective strategy on the rise in the BLM.

"We've had some success with planting firebreaks using green stripping. We're planting them with perennial grasses that green up later and stay green long after the cheatgrass is cured," said McCluskey.

The BLM has another tool to fight invasive plants. Use of chemicals such as pre-emergents may prevent cheatgrass from germinating, lessening its ability to outcompete natives.

McCluskey emphasized there is no one solution to control the spread of cheatgrass, or to improve habitats that have been invaded by cheatgrass.

"It's a combination of treatments," said McCluskey. "There is no panacea."

What does the future hold for wildlife habitats? McCluskey said it's important to look at what's happening on a very large scale.

"We're caught between a rock and a hard spot from our program perspective,” said McCluskey. “We're grappling internally with this in our agency. There's this Jeckyll and Hyde personality with a minerals mission on one side and a wildlife conservation mandate on the other side." That conflict makes land use allocation and management very challenging.

Understanding how sagebrush communities have changed over time and the management actions needed to restore these important habitats are keys to lessening the threat of invasive plants and restoring lands critical to mule deer survival.

Livestock management

Livestock management on western lands could be characterized as good, bad and ugly. Fortunately for mule deer, there’s a whole lot of good going on.

Utah Division of Wildlife Resources Big Game Coordinator Steve Cranney has seen elements of all three on lands in and around Utah and the West. But overall, Cranney is very positive about using cattle to manage wildlife habitat – if it’s done correctly.

“From our standpoint, livestock grazing has a lot of positives and negatives,” said Cranney. “When you do it right, it does have its use.”

Well-managed livestock grazing can improve the types and quantities of desirable plants, and maintain and create much-needed openings in dense habitat.

Cranney said his agency uses intense spring grazing on a number of wildlife management areas in big game winter ranges to graze grasses, and maintain and encourage growth of mule deer browse species.

“In the spring, cattle concentrate on the grass species, where the succulence is in the vegetation,” said Cranney. “Spring grazing encourages the growth of browse species such as sage and bitterbrush. The cattle are on the ground only for a month to a month and a half in the spring – strictly in the spring.”

Cranney said he varies how he uses cattle each year, depending on the status of the habitat, and the vegetation response he would like to see. He said his agency would use grazing even if it had the ability to use fire at any time because several plant species on deer winter range don’t respond favorably to fire, particularly sagebrush.

Cranney said, “We can’t just torch all winter range areas. Some browse species such as mountain mahogany and oakbrush respond favorably to fire, but sagebrush does not. How bitterbrush responds depends on the intensity of the fire. Spring grazing can be a valuable tool on many winter ranges.”

Cranney commented that spring grazing has other benefits, as well. It helps the local ranching community while helping mule deer.

“We enter into agreements with ranchers that help us,” said Cranney. “Spring grazing is very valuable to livestock people, too, because their cattle have been on hay all winter, and the ranchers are anxious to get their cattle off their ground so they can plant.”

Livestock grazing sounds like a win-win-win situation for state wildlife agencies, ranchers and mule deer. The bad and the ugly side come into play when livestock are not managed properly.

Poor livestock grazing practices can help spread invasive plants, interfere with plant succession, reduce nitrogen in the soil, and change the plant community. And improper livestock grazing in and around riparian areas may harm the stream and the rich diversity of wildlife that thrive in these environments. Overgrazing reduces water quality, changes stream flow, compacts and erodes soil, and affects native plants and animals that live alongside and in streams.

Tom Fleischner, in his 1994 Conservation Biology article, "Ecological costs of livestock grazing in western North America," said that livestock grazing has had "the most severe impact on one of the biologically richest habitats in the region," and states that, "much of the ecological integrity of a variety of North American habitats are at risk” because of poor grazing practices.

What kind of risk? Cottonwood/willow forests along arid western streams have declined about 90 percent since pre-settlement times. A 1988 report on "Restoring Degraded Riparian Areas on Western Rangelands" noted that "those narrow bands of green adjoining rivers, streams, and lakes, are crucial to the ecological health of arid western rangelands."

Cranney commented that cattle do the most harm in riparian areas.

“If they’re not fenced out, then they camp on it,” said Cranney. “The woody species and stream bank cover in riparian areas get taken out.” Cranney said this can be a serious problem, especially in states like Utah that are dry, and have limited riparian areas.

The good news is that the bad and ugly can be avoided. How can land managers manage livestock grazing for the benefit of people and wildlife? By establishing a sound range management program based on good range science and tailored to the local area. A good range management program should have the following elements:

1. Conduct prescribed burns to improve plant quality.

2. Do not graze stressed rangeland.

3. Control the number of livestock on rangeland to prevent overgrazing. Some ranchers recommend stocking at a rate less than 70 percent of average rainfall carrying capacity.

4. Use rotation grazing to prevent intensive spot grazing.

5. Fence riparian areas and provide off stream watering sources.

Cranney said state of the art wildlife management includes managing riparian areas as pastures with fence control.

“When the animals are in riparian areas, they are there strictly to benefit those areas,” commented Cranney. “The areas are grazed outside the fences.” This results in better grazing in upland areas, and minimal damage to streams and riparian habitat.

Grazing by livestock is a common and sometimes competing land use on many mule deer ranges. By Len Carpenter.Cranney said that sometimes the best wildlife management practice on winter ranges is not the most aesthetic, and visa versa.

“You go to Salt Lake City where cattle have been excluded for decades and it looks good from a watershed standpoint, very little open bare ground, and yet it’s poor for big game because there are few browse species,” said Cranney. “That’s why we concentrate on winter range areas. In most of Utah, the condition of winter range and the amount of it is critical.”

Cranney said one strategy his agency uses to protect mule deer winter range is wildlife easements.

Said Cranney, “Wildlife easements leave the property in the hands of the owners and allow them to conduct operations compatible with good wildlife management.” He also noted the most important aspect of wildlife easements is that they prevent subdivision of property into small ranchettes.

“Subdividing is the biggest enemy,” said Cranney.

Glenn Erickson, Chief of Montana Department of Fish, Wildlife and Parks Wildlife Management Bureau, echoed Cranney’s emphasis on keeping large tracts of land in private ownership. “We want to keep large, connected adjacent blocks of land in private ownership if we can.”

He said his agency places a strong emphasis on working with private landowners to improve livestock grazing practices.

“We’re providing consultation to landowners whenever they request it,” said Erickson. “We have a couple of people assigned full time to deal with grazing systems, and we have a couple of consultants that we work with through Montana State University. So there’s a big resource of information we provide.”

Erickson said his agency uses livestock grazing to improve the vegetation and soil on the state’s wildlife management areas.

“We modify how livestock graze, and where they graze,” said Erickson. “We typically try to protect riparian zones and manage vegetation zones in the pastures. It’s a rest rotation system, and the purpose is to benefit the vegetation for all species.”

Erickson commented that working with private landowners can multiply benefits to wildlife. “In some cases, we have our management area tied to adjacent private land, and we’re able to expand the advantage using a cooperative agreement,” said Erickson. “The landowner benefits and we benefit.”

Western wildlife agencies and provinces will continue to place an emphasis on positive working relationships with landowners and livestock managers to create mutually beneficial programs that ultimately enhance wildlife habitat for mule deer and other species. In doing so, land managers can assure that proper livestock management will continue to be a strong, positive change agent for mule deer habitat.


Mule Deer, Changing Landscapes, Changing Perspectives, is a series of non-technical articles based on technical papers from the book, “Mule Deer Conservation: Issues and Management Strategies” Published by The Berryman Institute, Utah State University.

The contents of this web page may be photocopied or reprinted for noncommercial purposes using the citation listed below:

Mule Deer Working Group. 2003. Mule Deer: Changing landscapes, changing perspectives. Mule Deer Working Group, Western Association of Fish and Wildlife Agencies.